Betty's life is in chaos after a series of very unfortunate events, but despite all that's happened, she's still in denial about one thing - how much Henry's departure has affected her. Meanwhile, Amanda turns to junk food and to Marc in dealing with the revelation that Fey Sommers was her real mother, and Wilhelmina schemes to turn the tragic Meade family events to her advantage.

Betty doesn't know what to do when she finds out Wilhelmina's hunky new bodyguard is taking the job to a whole new level. And then there's her own love life - now that Henry's back, can they just be friends? Meanwhile, Bradford uses Alexis' amnesia to re-write family history, Hilda comes to terms with recent events, and Amanda gets a surprising inheritance from Fey Sommers.

With Henry back in her life (sort of), Betty finds herself distracted by yet another guy - Gio, Mode's new sandwich guy, who both annoys and intrigues her. Meanwhile, the Mode-ites prepare for the social event of the season - the Meades' Black and White Ball, as Wilhelmina plots to get her wedding back on track, Amanda preps to make her first official appearance as Fey Sommer's illegitimate daughter, and fugitive-from-the law Claire invites herself to the party.

Betty takes steps toward becoming a writer and invests in a class that proves to be so challenging, she tosses her own writing assignment and instead submits a Mode story written by a woman who survived a bear attack - with disastrous results. Meanwhile Alexis, still suffering from amnesia, starts back at work with no memory of how to act like a girl; Daniel gets an ultimatum from their top advertiser - fire his tranny sister - and Amanda, on the hunt to find her dad's identity, learns a deep dark secret about Wilhelmina's past.

Much to Henry's distress, Betty tries internet dating and gets asked out on a bowling date. Meanwhile Marc finds himself attracted to someone who isn't his type, chubby but charming photographer Cliff. On the home front, Daniel and Alexis steel themselves for a "family" dinner with step-mom-to-be Wilhelmina, and Hilda, lost in her own world, doesn't notice that Justin is ditching school and running with the wrong crowd.

Betty and Henry continue their illicit romance and plan a secret date to "Wicked," but things don't go as planned when Daniel, thinking she's dating Gio, gifts her with tickets for the two of them instead - tickets which Gio refuses to give up. Meanwhile, Daniel woos an important advertiser, a lady with a penchant for younger men.

As Wilhelmina prepares for her big day, all is not quiet on the Mode front when she realizes her fabulous maid of honor, Victoria Beckham, is stealing her thunder. Meanwhile, Betty is wracked with guilt about her secret pact with Wilhelmina and about not telling Daniel that his future step-mom is dallying with her bodyguard. At the Suarez house, Ignacio prepares for his big day - becoming a U.S. citizen - but when Betty moves in with new love Henry, it puts a major strain on father/daughter relations.

In the wake of the disastrous Meade/Slater nuptials, Wilhelmina plots her next moves and Betty tries to make things right with Daniel. Meanwhile, Henry uses his charms to get the Mode weekend receptionist to let him in after hours so he can help Betty get something of vital importance.

A Wilhelmina-induced crisis at Mode finds everyone scrambling to re-create an issue on deadline. Desperate times call for desperate measures, as Daniel and Betty try to break wild child cover girl out of rehab for a re-shoot, and Alexis tries to charm the printer into extending their time. Meanwhile, Betty struggles with an unwelcome (and persistent) visitor who hopes to goad her into making good on a promise.

Henry reluctantly agrees to go on a double date with Betty and an unlikely couple - Hilda and Gio. Meanwhile, Wilhelmina's attempt to change her image as the queen of mean fails miserably after she tangles with America's most beloved golden girl.

Amanda visits a psychic for insight into her dad's identity. Meanwhile, disturbed about the message Mode is sending to Justin and his classmates after they visit the magazine, Betty pushes Daniel to use "real" models during fashion week; and Wilhelmina gets some unexpected news about her latest plans for the Meade dynasty.

Betty's new perfume is driving Henry wild and Betty crazy, literally, after she unwittingly begins using a deadly scent concocted by the late Fey Sommers and starts acting very strangely. Meanwhile, Meade matriarch Claire goes on the stand for Fey's murder, a vindictive Amanda withholds important information, and Wilhelmina takes steps to become a member of the Meade clan - one way or the other.

Betty gets an assignment from Daniel to interview "Phil Roth," thinking it's a sit down with the great novelist, but later learning this Phil Roth is a writer who has penned a series of books about picking up women. Meanwhile Daniel hooks up with a sexy new lady, Renee, only to learn they have a shocking connection to each other, and Amanda and Marc hatch a plan to reach out to the man they believe is Amanda's biological dad.